I've figured out that these are Kluson 538 tuners from the late 60's/early 70's, and likely taken off of a Gibson bass. The more I play this H22 the more I suspect that it belonged, for many years, to a working bassist. Things like the intonation and action being setup so well and the cheesy Harmony tuners being replaced with what, at that time, were high quality tuners (apparently these old Klusons still have resale value). I may just keep these elephant ears on there out of respect for the previous owner, whoever he/she was.