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Recommend a fretless...

The "mwaaahh" is from roundwound strings. The Jaco tone is a combination of that and the bridge pickup on a Jazz Bass. Put some flats on and don't favor the bridge pickup and you have a nice fretless sound. Any fretless bass, even Jaco's bass in his hands, can give this kind of sound.
The tone I'm looking for in a fretles is the really deep, kinda growly, almost uprightish tone, and I'm wondering what kind of fretless would get me that for pretty cheap. I'm talkin about in the $400-600 range.
If by "uprightish" you mean quick-sustaining "plunk" you'll want an acoustic or acoustic-style bass. Otherwise, in the 400-600 range there are about ten zillion great options.

My first fretless was a $450 Dean Edge, it was fantastic. Being the first generation of Edge, it had a passive jazz bass style EQ with two EMG-HZ humbucking soapbars. GREAT sound.
 
I own a Rob Allen Deep 4 which I suspect is close to the tone you want (with LaBella Deep Talkin' flats ... strings are a major contributor to fretless sound). This is uprightish, but with more sustain. To back off on the sustain, you can go with Rotosound88s. But I think this may (at $4500) be out of your price range.

I also own a Dean Edge Fretless 4 ($300 purchase) with some upgrades (Badass II bridge; DC4 pickups; Aguilar OBP-1) that improve the sound (the original sound was adequate). I have heavy GHS Boomers on this, which give very little 'mwahh' (I think because they are heavy) but more a Fenderish fretted tone.

I like fretless a lot, because I feel a lot freeer on the fretboard ... less mechanical.
 
Not being a real J style bass player ... well normally anyway, and I do own one, I dig the Lakland JO fretless. I have also owned a 70s fretless P (not to my liking) I really think my Lakland JO fretless is the bomb! I have previously owned a Pedulla and a Modulus, but the rosewood board Lakland wins hands down! It's all in the setup.



:) ... That said ... I can see that a fretlless P style bass with a J taper lined rosewood neck XL chromes flats and a Darkstar pickup may be "THE" fretless to own! ... note to self call Dan Lakin.
 
Have you thought about defretting that Ibanez you're selling? Just a thought - it's hollowbody, has a sweet-spot pickup, you could easily fit felt or sponge mutes if that's what you're after... If I had one and was interested in thuddy fretless, I'd be very tempted. And it'd cost you next to nothing except a bit of time and a radiused sanding block from stewmac.

Even though it's just about everything you've said or been advised here you don't want, I've gotten compliments from my brother on the upright-ness of my Ibanez ATK defret, which is strung with Fender flats. Of course, it has a bridge pickup only, and active EQ, so make of that what you will. I have to say that flats make a huge difference. I have played a few fretli with rounds, and I much prefer the sound and feel of flats on fretless. I use fretteds if I want edge and clank. Fretless is more about thud for me (no offense, Jaco).

Good luck,
Steve