It says that the DC voltage is low there. NOW, we need to identify WHY.
If you remove the two tubes on that power amp PCB, the voltage after the variable stage should rise to approx. what it's supposed to be. If it doesn't, it indicates a problem with the variable plate voltage circuit.
If it does, it doesn't mean that it's not the plate voltage control circuit, BUT it could also be a problem with either tube or the resistors or C3 surrounding the tubes. With the tubes out of the circuit (and the power disconnected), wait a few minutes, verify that the power supply is fully discharged, you should be able to measure most if not all in circuit.
If they all check out, and replacing the tubes (swapping with known good tubes) doesn't change things, it points to the plate voltage control circuit. If that's the case, I would expect that C5 is leaky, pulling down the base control voltage on the pass element. That said, if any of those resistors are way off spec, that too could cause the same symptoms.
C5 needs to be 47uF/450V because the initial conditions can be higher than the steady state conditions.