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Nice Work.
The definition of “growl” varies by person. @Bruce Johnson did an excellent job describing what is basically controlled fret buzz, used by a lot of well known players. There’s also a version of “growl” referred to by many that is not derived from fret buzz only, and is much more related to tone / harmonic content. It’s not exactly distortion, but could be close to distortion in a specific frequency range. This version of growl generally includes the term “low-mids” somewhere in the discussion & description. Some classic P-Bass tones are often described as having “growl”. It can also be related to specific amp / speaker combinations (example; Ampeg SVT & 810 cab) and in this case is very much likely to be distortion and/or certain harmonics that stand out above others.
I think you are referring to the growl related to low-mids & related harmonics.
Listen to this beginning at 3:30 for an example of tone related growl.
Actually, I don’t even know *** “growl” even is in relation to Warwicks. I know Jameson didn’t growl (flats, high action) but Entwhistle did (rounds, fret buzz). Clearly I’m missing the point of this thread, or “growl” needs to be more explicitly described (for me, anyway).

Fret buzz and growl? 100% unrelted!
it's the pickup+pickup placement+Bright strings+plucking closer to the Bridge
There seem to be different ideas about what growl is. To me it is when the lower notes have lots of overtones in them, as opposed to just the fundamentals. This is a factor of pickups, their placement, and electronics. I got my Gary Willis bass because of how it sounds. The pickup is close to the bridge and the electronics accentuate the mids which makes the low B and E sound very rich in harmonics.
My Am. Ultra PJ growls well, I believe because of the rather substantial bridge pickup.So warwicks are known for their "growl" (especially the thumb), but I want to know is what causes that sound, and why it doesn't sound, say like a jazz bass, I mean they've got the same pickups, what separates a Thumb BO4 from an American Professional Jazz.
My definition of growl is not exactly fret buzz.
Fret buzz. Bruce Thomas has this in spades. His Precision sounds like it’s about to fizz out. Attitude for days.
ding ding dingThe definition of “growl” varies by person.