Interesting, never knew that. But that doesn't change the equation all the more reason for the bass to be in tune with it.
But here again, the limitations of the instrument come into play:
A very long scale, fretless instruments. If you' re not thoroughly schooled on your neck hand, maybe you're off a pinch.
Straight bridge, no adjusting individually the length for intonation.
Really high action, possibly sharp even if you're dead on a note.
And it can be bowed.
You can think of a lot of ways to be out of dead-on-the-money in tune, and often, everything's out just a hair.
This stuff can make you crazy, and utterly torture the rare person with true perfect pitch:
I once was in the opening act for an outdoor festival gig where Ronnie Milsap was the top of the bill. His rider required at least a seven foot Yamaha or Steinway, tuned at its original location, retuned once delivered to the venue and double checked no more than two hours before he sat down to play it.
It was all done and parked Stage Right for the opening acts. When it was time to move it center stage for his show, rolling it on that big Y dolly with the go-kart wheels was not allowed. About 29 of us got all around it, picked it straight up, walked it about 15 feet, and set it down as gingerly as a live bomb.
Luckily, it was just fine.