Since this thread came up, I've been trying to think of how I can put into words the "feel" of tube amps that explains how I can tell when I'm playing them. I think I finally came up with it.
1. People automatically think "tube = compression" but I don't hear them that way. Sure, when you crank them they're compressing, but when you run them clean, I don't know if the signal is technically being compressed by tubes or not, but it doesn't sound like it to me. They actually sound to me the least compressed of any amps I've ever played as long as I'm running them with decent headroom. No matter how hard I dig in, they dig in with me. No matter how softly I play, they go that much softer. I don't get that dynamic response in solid state designs. They always seem to have a point where they don't follow my dynamics that closely. Some do a really good job with it, but none pick up on the most subtle nuances of it like a clean running tube amp does.
2. I don't think the sheer physicality of your signal traveling through giant iron transformers and a network of glass-enclosed vacuum tubes can be taken for granted. All those little wires getting heated up, all that glass getting heated up, your bass signal traveling up and down that network of iron and glass until it hits the speakers...you can feel it when you play the bass or guitar and you can hear it when you've educated your ear to the difference. The emulations do very good at duplicating those sounds, but I think that physical reaction just can't be duplicated.
But one day, I do believe Seamonkey will be right about his claims that SS technology can emulate everything about tubes. I never thought they could even come close, and it's only been in the last few years that I think they have. But I've heard recordings of them and I'm hard pressed to tell the difference whereas I always believed I could in the past. The emulations are quite good now, and it's only a matter of time before it can all be duplicated, feel and all. But that's how I can best describe the "feel" differences I notice when I play a tube amp vs a solid state emulation, and that's why I don't think it's there yet.